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Atrociraptor

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Atrociraptor
Temporal range: Late Cretaceous (Maastrichtian), 68.5 Ma
Holotype jaw remains at the Royal Tyrrell Museum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Family: Dromaeosauridae
Clade: Eudromaeosauria
Subfamily: Saurornitholestinae
Genus: Atrociraptor
Currie & Varricchio, 2004
Species:
A. marshalli
Binomial name
Atrociraptor marshalli
Currie & Varricchio, 2004

Atrociraptor (meaning "savage thief"[1]) is a genus of saurornitholestine dromaeosaurid theropod dinosaur from the Late Cretaceous (Maastrichtian stage) of Alberta, Canada.

Discovery[edit]

Skeletal diagram showing the position of the holotype jaw remains

In 1995, Wayne Marshall discovered a partial dromaeosaur skull in the Horseshoe Canyon Formation of Drumheller, Alberta, about 5 km west of the Royal Tyrrell Museum of Palaeontology. Jaw fragments and teeth on the hillside led to the discovery of the specimen.In 2004 Philip J. Currie and David Varricchio named and described the type species of Atrociraptor: Atrociraptor marshalli. The generic name is derived from the Latin atrox, "savage", and raptor, "seizer". The specific name honours Marshall.[1]

The holotype specimen of Atrociraptor, RTMP 95.166.1, was discovered by in 1995, The only known specimen consists of parts of the upper and lower jaws—both premaxillae, a right maxilla, both dentaries—teeth and numerous small fragments. The skull appears to have been unusually short and tall. The teeth are relatively straight, but they emerge from the tooth sockets at an angle to the jaw line, resulting in a strongly raked row of teeth. A number of isolated teeth (previously referred to Saurornitholestes) have also been recovered from the Horseshoe Canyon Formation;[2] they can be recognized by their unusually large serrations.

Description[edit]

Size compared to a human

Atrociraptor was a relatively small dromaeosaur, measuring 2 m (6.6 ft) long and weighing 15 kg (33 lb).[3] It differs from Bambiraptor and other Dromaeosaurids in its more isodont dentition—the teeth have different sizes but the same form—and short deep snout. A skull opening, the maxillary fenestra, is relatively large and positioned right above another opening, the promaxillary fenestra, a condition not known from other species.

Classification[edit]

Hypothetical life restoration

Atrociraptor was by its describers assigned to the Velociraptorinae within a larger Dromaeosauridae. However, in 2009 Currie published a cladistic analysis showing Atrociraptor to be a member of the Saurornitholestinae.[4] A 2022 study and analysis further vindicates this position and classifies Atrociraptor as a derived saurornitholestine, along with Acheroraptor.[5][6][7][8]

The cladogram below shows the position of Atrociraptor within Dromaeosauridae according to the 2022 analysis by Jasinski and colleagues:[9]

Dromaeosauridae

Palaeoenvironment[edit]

The holotype was found in layers also containing an Albertosaurus bonebed. This bonebed is located at the top of Unit 4 of the Horseshoe Canyon Formation,[10] which dates to about 68.5 million years ago.[11]

See also[edit]

References[edit]

  1. ^ a b Currie, P. J. and D. J. Varricchio (2004). "A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada". Pp. 112–132 in P. J. Currie, E. B. Koppelhus, M. A. Shugar and J. L. Wright. (eds.), Feathered Dragons. Indianapolis: Indiana University Press. [1]
  2. ^ Ryan, M. J., P. J. Currie, et al. (1998). "Baby hadrosaurid material associated with an unusually high abundance of Troodon teeth from the Horseshoe Canyon Formation, Upper Cretaceous, Alberta, Canada". Gaia 16: 123–133
  3. ^ Paul, G.S., 2010, The Princeton Field Guide to Dinosaurs, Princeton University Press p. 136
  4. ^ N.R. Longrich and P.J. Currie, 2009, "A microraptorine (Dinosauria-Dromaeosauridae) from the Late Cretaceous of North America". Proceedings of the National Academy of Sciences 106: 5002-5007
  5. ^ Powers, Mark J.; Fabbri, Matteo; Doschak, Michael R.; Bhullar, Bhart-Anjan S.; Evans, David C.; Norell, Mark A.; Currie, Philip J. (2022). "A new hypothesis of eudromaeosaurian evolution: CT scans assist in testing and constructing morphological characters". Journal of Vertebrate Paleontology. 41 (5): e2010087. doi:10.1080/02724634.2021.2010087. S2CID 247039404.
  6. ^ Chiarenza, Alfio Alessandro; Fiorillo, Anthony R.; Tykoski, Ronald S.; McCarthy, Paul J.; Flaig, Peter P.; Contreras, Dori L. (2020). "The first juvenile dromaeosaurid (Dinosauria: Theropoda) from Arctic Alaska". PLOS ONE. 15 (7): e0235078. Bibcode:2020PLoSO..1535078C. doi:10.1371/journal.pone.0235078. PMC 7343144. PMID 32639990.
  7. ^ Pittman, Michael; Xu, Xing (2020). "Pennaraptoran Theropod Dinosaurs Past Progress and New Frontiers". Bulletin of the American Museum of Natural History. 440 (1): 1. doi:10.1206/0003-0090.440.1.1.
  8. ^ Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019). "A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight". PeerJ. 7: e7247. doi:10.7717/peerj.7247. PMC 6626525. PMID 31333906.
  9. ^ Jasinski, Steven E.; Sullivan, Robert M.; Carter, Aja M.; Johnson, Erynn H.; Dalman, Sebastian G.; Zariwala, Juned; Currie, Philip J. (2022-11-07). "Osteology and reassessment of Dineobellator notohesperus , a southern eudromaeosaur (Theropoda: Dromaeosauridae: Eudromaeosauria) from the latest Cretaceous of New Mexico". The Anatomical Record. 306 (7): 1712–1756. doi:10.1002/ar.25103. ISSN 1932-8486. PMID 36342817. S2CID 253382718.
  10. ^ Larson, D. W., Brinkman, D. B., & Bell, P. R. (2010). Faunal assemblages from the upper Horseshoe Canyon Formation, an early Maastrichtian cool-climate assemblage from Alberta, with special reference to the Albertosaurus sarcophagus bonebed This article is one of a series of papers published in this Special Issue on the theme Albertosaurus. Canadian Journal of Earth Sciences, 47(9), 1159-1181.
  11. ^ Arbour, Victoria (2010). "A Cretaceous armoury: Multiple ankylosaurid taxa in the Late Cretaceous of Alberta, Canada and Montana, USA". Journal of Vertebrate Paleontology. 30 (Supplement 2): 55A. doi:10.1080/02724634.2010.10411819. S2CID 220429286.